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  Animalia
      Arthropoda
            Insecta
                    Coleoptera
                            Ceratocanthidae
                                     Perignamptus
                                             Perignamptus rossi
                      Polyphaga
                        Scarabeiformia
                          Scarabaeoidea
                            Hybosoridae
                              Ceratocanthinae
                                     [Ceratocanthidae]
                                Ceratocanthini
                                     Ceratocanthus
                                           [Acanthocerus]
                                           [Gymnoropterus]
                                           [Sphaeromorphus]
                                             Ceratocanthus aeneus
                                                   [Sphaeromorphus volvox]
                                     Germarostes
                                       Germarostes (Germarostes)
                                             Germarostes aphodioides
                                             Germarostes globosus
                                     Philharmostes
                                             Philharmostes apterus
                                     Cryptosphaeroides
                                             Cryptosphaeroides ankaranensis
                                             Cryptosphaeroides hippocrepis
                                             Cryptosphaeroides tenrec
                                     Pseudosynarmostes
                                             Pseudosynarmostes mitsinjo

Problematic Rooting (148933)
                                             Acanthocerus crucifer
                                                   [Acanthocerus cruciger]
                                             Acanthocerus laevistriatus
                                             Acanthocerus lobatus
                                             Acanthocerus tuberculatus