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  Animalia
      Arthropoda
            Arachnida
                    Pseudoscorpiones
                           [Diplosphyronida]
                           [Heterosphyronida]
                           [Monosphyronida]
                           [Pseudoscorpionida]
                      Epiocheirata
                          Chthonioidea
                            Chthoniidae
                                   [Chthonidae]
                              Chthoniinae
                                     Anisoditha
                                             Anisoditha curvidigitata
                                     Aphrastochthonius
                                             Aphrastochthonius alteriae
                                             Aphrastochthonius cubanus
                                             Aphrastochthonius grubbsi
                                             Aphrastochthonius major
                                             Aphrastochthonius pachysetus
                                             Aphrastochthonius palmitensis
                                             Aphrastochthonius parvus
                                             Aphrastochthonius patei
                                             Aphrastochthonius pecki
                                             Aphrastochthonius russelli
                                             Aphrastochthonius similis
                                             Aphrastochthonius tenax
                                             Aphrastochthonius verapazanus
                                     Apochthonius
                                             Apochthonius coecus
                                             Apochthonius colecampi
                                             Apochthonius diabolus
                                             Apochthonius forbesi
                                             Apochthonius grubbsi
                                             Apochthonius hobbsi
                                             Apochthonius holsingeri
                                             Apochthonius hypogeus
                                             Apochthonius indianensis
                                             Apochthonius intermedius
                                             Apochthonius irwini
                                             Apochthonius knowltoni
                                             Apochthonius magnanimus
                                             Apochthonius malheuri
                                             Apochthonius maximus
                                             Apochthonius minimus
                                             Apochthonius minor
                                             Apochthonius moestus
                                             Apochthonius mysterius
                                             Apochthonius occidentalis
                                             Apochthonius paucispinosus
                                             Apochthonius russelli
                                             Apochthonius titanicus
                                             Apochthonius typhlus
                                     Austrochthonius
                                           [Cecoditha]
                                           [Paraustrochthonius]
                                             Austrochthonius argentinae
                                             Austrochthonius australis
                                             Austrochthonius bolivianus
                                             Austrochthonius cavicola
                                             Austrochthonius chilensis
                                               Austrochthonius chilensis chilensis
                                               Austrochthonius chilensis magalhanicus
                                               Austrochthonius chilensis transversus
                                             Austrochthonius easti
                                             Austrochthonius iguazuensis
                                             Austrochthonius insularis
                                             Austrochthonius mordax
                                             Austrochthonius muchmorei
                                             Austrochthonius paraguayensis
                                             Austrochthonius parvus
                                                   [Cecoditha parva]
                                             Austrochthonius persimilis
                                             Austrochthonius rapax
                                             Austrochthonius semiserratus
                                             Austrochthonius strigosus
                                             Austrochthonius tullgreni
                                             Austrochthonius zealandicus
                                               Austrochthonius zealandicus obscurus
                                               Austrochthonius zealandicus zealandicus
                                     Caribchthonius
                                           [Caribohthonius]
                                             Caribchthonius butleri
                                             Caribchthonius orthodentatus
                                     Chiliochthonius
                                             Chiliochthonius centralis
                                             Chiliochthonius montanus
                                     Chthonius
                                           [Kewochthonius]
                                       Chthonius (Chthonius)
                                             Chthonius absoloni
                                             Chthonius agazzii
                                             Chthonius alpicola
                                             Chthonius apollinis
                                             Chthonius azerbaidzhanus
                                             Chthonius balazuci
                                             Chthonius bogovinae
                                               Chthonius bogovinae bogovinae
                                               Chthonius bogovinae latidentatus
                                             Chthonius brandmayri
                                             Chthonius caoduroi
                                             Chthonius caprai
                                             Chthonius carinthiacus
                                             Chthonius cavernarum
                                             Chthonius cavophilus
                                             Chthonius cebenicus
                                             Chthonius cephalotes
                                             Chthonius chamberlini
                                             Chthonius comottii
                                             Chthonius cryptus
                                             Chthonius dacnodes
                                             Chthonius dalmatinus
                                             Chthonius decoui
                                             Chthonius densedentatus
                                             Chthonius diophthalmus
                                             Chthonius doderoi
                                               Chthonius doderoi doderoi
                                               Chthonius doderoi horridus
                                             Chthonius ellingseni
                                             Chthonius elongatus
                                             Chthonius euganeus
                                             Chthonius exarmatus
                                             Chthonius graecus
                                             Chthonius guglielmii
                                             Chthonius halberti
                                             Chthonius herbarii
                                             Chthonius heterodactylus
                                             Chthonius heurtaultae
                                             Chthonius hungaricus
                                             Chthonius ilvensis
                                             Chthonius imperator
                                             Chthonius ischnocheles
                                               Chthonius ischnocheles ischnocheles
                                               Chthonius ischnocheles reductus
                                             Chthonius ischnocheloides
                                             Chthonius italicus
                                             Chthonius iugoslavicus
                                             Chthonius jalzici
                                             Chthonius jonicus
                                             Chthonius jugorum
                                             Chthonius karamanianus
                                             Chthonius lanzai
                                               Chthonius lanzai lanzai
                                               Chthonius lanzai vannii
                                             Chthonius leoi
                                             Chthonius leruthi
                                             Chthonius lesnik
                                             Chthonius lessiniensis
                                             Chthonius ligusticus
                                               Chthonius ligusticus brevis
                                               Chthonius ligusticus ligusticus
                                             Chthonius lindbergi
                                             Chthonius litoralis
                                             Chthonius lucifugus
                                             Chthonius macedonicus
                                             Chthonius magnificus
                                             Chthonius malatestai
                                             Chthonius mauritanicus
                                             Chthonius mayi
                                             Chthonius mazaurici
                                               Chthonius mazaurici coironi
                                               Chthonius mazaurici mazaurici
                                             Chthonius microphthalmus
                                             Chthonius mingazzinii
                                             Chthonius minotaurus
                                             Chthonius monicae
                                             Chthonius motasi
                                             Chthonius multidentatus
                                             Chthonius occultus
                                             Chthonius ognjankae
                                             Chthonius ohridanus
                                             Chthonius orthodactyloides
                                             Chthonius orthodactylus
                                             Chthonius pacificus
                                             Chthonius paganus
                                             Chthonius paludis
                                             Chthonius paolettii
                                             Chthonius persimilis
                                             Chthonius petrochilosi
                                             Chthonius pivai
                                             Chthonius ponticoides
                                             Chthonius ponticus
                                             Chthonius porevidi
                                             Chthonius prove
                                             Chthonius pusillus
                                             Chthonius pygmaeus
                                             Chthonius radigost
                                             Chthonius radjai
                                             Chthonius raridentatus
                                             Chthonius ressli
                                             Chthonius rhodochelatus
                                             Chthonius ruffoi
                                             Chthonius satapliaensis
                                             Chthonius sestasi
                                             Chthonius shelkovnikovi
                                             Chthonius shulovi
                                             Chthonius stammeri
                                             Chthonius stevanovici
                                             Chthonius strinatii
                                             Chthonius submontanus
                                             Chthonius subterraneus
                                               Chthonius subterraneus meuseli
                                               Chthonius subterraneus subterraneus
                                             Chthonius tadzhikistanicus
                                             Chthonius tenuis
                                             Chthonius thessalus
                                             Chthonius trebinjensis
                                             Chthonius troglobius
                                             Chthonius troglodites
                                             Chthonius tzanoudakisi
                                             Chthonius vodan
                                             Chthonius zmaj
                                             Chthonius (Chthonius) ischnocheles
                                             Chthonius (Chthonius) orthodactylum
                                             Chthonius (Chthonius) pacificus
                                             Chthonius (Chthonius) paganus
                                             Chthonius (Chthonius) paludis
                                       Chthonius (Ephippiochthonius)
                                             Chthonius aegatensis
                                             Chthonius aguileraorum
                                             Chthonius amatei
                                             Chthonius anatolicus
                                             Chthonius anophthalmus
                                             Chthonius apulicus
                                             Chthonius asturiensis
                                             Chthonius atlantis
                                             Chthonius balearicus
                                             Chthonius bartolii
                                             Chthonius bauneensis
                                             Chthonius bellesi
                                             Chthonius berninii
                                             Chthonius bidentatus
                                             Chthonius boldorii
                                             Chthonius bolivari
                                             Chthonius cabreriensis
                                             Chthonius cassolai
                                             Chthonius catalonicus
                                             Chthonius cavicola
                                             Chthonius chius
                                             Chthonius concii
                                             Chthonius corcyraeus
                                             Chthonius corsicus
                                             Chthonius creticus
                                             Chthonius daedaleus
                                             Chthonius distinguendus
                                             Chthonius dubius
                                             Chthonius elbanus
                                             Chthonius fuscimanus
                                             Chthonius gasparoi
                                             Chthonius genuensis
                                             Chthonius gestroi
                                             Chthonius gibbus
                                             Chthonius girgentiensis
                                             Chthonius giustii
                                             Chthonius gracilimanus
                                             Chthonius grafittii
                                             Chthonius hiberus
                                             Chthonius hispanus
                                             Chthonius insularis
                                             Chthonius iranicus
                                             Chthonius kabylicus
                                             Chthonius kemza
                                             Chthonius kewi
                                             Chthonius kupalo
                                             Chthonius longesetosus
                                             Chthonius lucanus
                                             Chthonius lychnidis
                                             Chthonius machadoi
                                               Chthonius machadoi canariensis
                                               Chthonius machadoi machadoi
                                             Chthonius mahnerti
                                             Chthonius maltensis
                                             Chthonius mariolae
                                             Chthonius maroccanus
                                             Chthonius mayorali
                                             Chthonius microtuberculatus
                                             Chthonius minous
                                               Chthonius minous minous
                                               Chthonius minous peramae
                                             Chthonius minutus
                                             Chthonius nanus
                                             Chthonius nerjaensis
                                             Chthonius nidicola
                                             Chthonius nudipes
                                             Chthonius parmensis
                                             Chthonius pieltaini
                                             Chthonius pinai
                                             Chthonius platakisi
                                             Chthonius poeninus
                                             Chthonius ponsi
                                             Chthonius poseidonis
                                             Chthonius pyrenaicus
                                             Chthonius remyi
                                             Chthonius rimicola
                                             Chthonius romanicus
                                             Chthonius ruizporteroi
                                             Chthonius sacer
                                             Chthonius samius
                                             Chthonius schmalfussi
                                             Chthonius scythicus
                                             Chthonius sendrai
                                             Chthonius serbicus
                                             Chthonius setosus
                                             Chthonius siculus
                                             Chthonius siscoensis
                                             Chthonius tetrachelatus
                                             Chthonius troglophilus
                                             Chthonius tuberculatus
                                             Chthonius vachoni
                                             Chthonius ventalloi
                                               Chthonius ventalloi cazorlensis
                                               Chthonius ventalloi ventalloi
                                             Chthonius verai
                                             Chthonius vid
                                             Chthonius virginicus
                                             Chthonius zoiai
                                             Chthonius (Ephippiochthonius) tetrachelatus
                                             Chthonius (Ephippiochthonius) virginicus
                                       Chthonius (Globochthonius)
                                             Chthonius abnormis
                                             Chthonius caligatus
                                             Chthonius cavernicola
                                             Chthonius cerberus
                                             Chthonius globifer
                                             Chthonius pancici
                                             Chthonius perun
                                             Chthonius polychaetus
                                             Chthonius purgo
                                             Chthonius simplex
                                             Chthonius spelaeophilus
                                               Chthonius spelaeophilus histricus
                                               Chthonius spelaeophilus spelaeophilus
                                             Chthonius vandeli
                                       Chthonius (Hesperochthonius)
                                             Chthonius californicus
                                             Chthonius oregonicus
                                             Chthonius spingolus
                                             Chthonius (Hesperochthonius) californicus
                                             Chthonius (Hesperochthonius) oregonicus
                                             Chthonius (Hesperochthonius) spingonicus
                                       Chthonius (Microchthonius)
                                             [Microchthonius]
                                             Chthonius karamani
                                                   [Microchthonius karamani]
                                             Chthonius rogatus
                                             Chthonius dilatimanus
                                             Chthonius johnstoni
                                             Chthonius spingonicus
                                     Congochthonius
                                             Congochthonius nanus
                                     Drepanochthonius
                                             Drepanochthonius horridus
                                     Francochthonius
                                             Francochthonius hirsutus
                                     Kleptochthonius
                                           [Heterochthonius]
                                       Kleptochthonius (Chamberlinochthonius)
                                             Kleptochthonius affinis
                                             Kleptochthonius anophthalmus
                                             Kleptochthonius attenuatus
                                             Kleptochthonius barri
                                             Kleptochthonius binoculatus
                                             Kleptochthonius cerberus
                                             Kleptochthonius charon
                                             Kleptochthonius daemonius
                                             Kleptochthonius erebicus
                                             Kleptochthonius gertschi
                                             Kleptochthonius hageni
                                             Kleptochthonius henroti
                                             Kleptochthonius hetricki
                                             Kleptochthonius hubrichti
                                             Kleptochthonius infernalis
                                             Kleptochthonius krekeleri
                                             Kleptochthonius lutzi
                                             Kleptochthonius microphthalmus
                                             Kleptochthonius myopius
                                             Kleptochthonius orpheus
                                             Kleptochthonius packardi
                                             Kleptochthonius pluto
                                             Kleptochthonius proserpinae
                                             Kleptochthonius proximosetus
                                             Kleptochthonius regulus
                                             Kleptochthonius rex
                                             Kleptochthonius similis
                                             Kleptochthonius stygius
                                             Kleptochthonius tantalus
                                             Kleptochthonius (Chamberlinochthonius) affinis
                                             Kleptochthonius (Chamberlinochthonius) anophtha
                                             Kleptochthonius (Chamberlinochthonius) attenuat
                                             Kleptochthonius (Chamberlinochthonius) barri
                                             Kleptochthonius (Chamberlinochthonius) binocula
                                             Kleptochthonius (Chamberlinochthonius) cerberus
                                             Kleptochthonius (Chamberlinochthonius) charon
                                             Kleptochthonius (Chamberlinochthonius) daemoniu
                                             Kleptochthonius (Chamberlinochthonius) erebicus
                                             Kleptochthonius (Chamberlinochthonius) gertschi
                                             Kleptochthonius (Chamberlinochthonius) hageni
                                             Kleptochthonius (Chamberlinochthonius) henroti
                                             Kleptochthonius (Chamberlinochthonius) hetricki
                                             Kleptochthonius (Chamberlinochthonius) hubricht
                                             Kleptochthonius (Chamberlinochthonius) infernal
                                             Kleptochthonius (Chamberlinochthonius) kerkeler
                                             Kleptochthonius (Chamberlinochthonius) lutzi
                                             Kleptochthonius (Chamberlinochthonius) micropht
                                             Kleptochthonius (Chamberlinochthonius) myopius
                                             Kleptochthonius (Chamberlinochthonius) orpheus
                                             Kleptochthonius (Chamberlinochthonius) packardi
                                             Kleptochthonius (Chamberlinochthonius) pluto
                                             Kleptochthonius (Chamberlinochthonius) proserpi
                                             Kleptochthonius (Chamberlinochthonius) proximos
                                             Kleptochthonius (Chamberlinochthonius) regulus
                                             Kleptochthonius (Chamberlinochthonius) rex
                                             Kleptochthonius (Chamberlinochthonius) similis
                                             Kleptochthonius (Chamberlinochthonius) stygius
                                             Kleptochthonius (Chamberlinochthonius) tantalus
                                             Kleptochthonius (Kleptochitonius) geophilus
                                             Kleptochthonius (Kleptochitonius) magnus
                                             Kleptochthonius (Kleptochitonius) multispinosus
                                             Kleptochthonius (Kleptochitonius) oregonus
                                       Kleptochthonius (Kleptochthonius)
                                             Kleptochthonius crosbyi
                                             Kleptochthonius geophilus
                                             Kleptochthonius inusitatus
                                             Kleptochthonius magnus
                                             Kleptochthonius multispinosus
                                             Kleptochthonius oregonus
                                             Kleptochthonius polychaetus
                                             Kleptochthonius sheari
                                             Kleptochthonius griseomanus
                                             Kleptochthonius lewisorum
                                     Lagynochthonius
                                             Lagynochthonius annamensis
                                             Lagynochthonius arctus
                                             Lagynochthonius australicus
                                             Lagynochthonius bakeri
                                             Lagynochthonius brincki
                                             Lagynochthonius callidus
                                             Lagynochthonius cavicola
                                             Lagynochthonius curvidigitatus
                                             Lagynochthonius dybasi
                                             Lagynochthonius exiguus
                                             Lagynochthonius ferox
                                             Lagynochthonius flavus
                                             Lagynochthonius gigas
                                             Lagynochthonius guasirih
                                             Lagynochthonius hamatus
                                             Lagynochthonius himalayensis
                                             Lagynochthonius hygricus
                                             Lagynochthonius indicus
                                             Lagynochthonius innoxius
                                             Lagynochthonius insulanus
                                             Lagynochthonius irmleri
                                             Lagynochthonius johni
                                             Lagynochthonius kapi
                                             Lagynochthonius kenyensis
                                             Lagynochthonius minor
                                             Lagynochthonius mordor
                                             Lagynochthonius nagaminei
                                             Lagynochthonius novaeguineae
                                             Lagynochthonius paucedentatus
                                             Lagynochthonius ponapensis
                                             Lagynochthonius proximus
                                             Lagynochthonius pugnax
                                             Lagynochthonius roeweri
                                             Lagynochthonius salomonensis
                                             Lagynochthonius sinensis
                                             Lagynochthonius thorntoni
                                             Lagynochthonius tongae
                                             Lagynochthonius tonkinensis
                                             Lagynochthonius typhlus
                                             Lagynochthonius zicsii
                                     Malcolmochthonius
                                             Malcolmochthonius malcolmi
                                             Malcolmochthonius oregonus
                                             Malcolmochthonius perplexus
                                     Maorichthonius
                                             Maorichthonius mortenseni
                                     Mexichthonius
                                             Mexichthonius exoticus
                                             Mexichthonius pacal
                                             Mexichthonius unicus
                                     Mundochthonius
                                             Mundochthonius alpinus
                                             Mundochthonius asiaticus
                                             Mundochthonius basarukini
                                             Mundochthonius bifurcatus
                                             Mundochthonius carpaticus
                                             Mundochthonius cavernicola
                                             Mundochthonius decoui
                                             Mundochthonius dominicanus
                                             Mundochthonius erosidens
                                             Mundochthonius gallaecicus
                                             Mundochthonius holsingeri
                                             Mundochthonius japonicus
                                               Mundochthonius japonicus imadatei
                                               Mundochthonius japonicus japonicus
                                               Mundochthonius japonicus scolytidis
                                               Mundochthonius japonicus tripartitus
                                             Mundochthonius kiyoshii
                                             Mundochthonius magnus
                                             Mundochthonius mexicanus
                                             Mundochthonius minusculus
                                             Mundochthonius montanus
                                             Mundochthonius pacificus
                                             Mundochthonius rossi
                                             Mundochthonius sandersoni
                                             Mundochthonius singularis
                                             Mundochthonius styriacus
                                             Mundochthonius ussuricus
                                     Neochthonius
                                             Neochthonius amplus
                                             Neochthonius imperialis
                                             Neochthonius latisetosus
                                             Neochthonius stanfordianus
                                             Neochthonius troglodytes
                                     Paraliochthonius
                                           [Morikawia]
                                             Paraliochthonius azanius
                                             Paraliochthonius canariensis
                                             Paraliochthonius carpenteri
                                             Paraliochthonius cavalensis
                                             Paraliochthonius hoestlandti
                                             Paraliochthonius insulae
                                             Paraliochthonius johnstoni
                                             Paraliochthonius martini
                                             Paraliochthonius mirus
                                             Paraliochthonius puertoricensis
                                             Paraliochthonius singularis
                                             Paraliochthonius takashimai
                                             Paraliochthonius tenebrarum
                                             Paraliochthonius weygoldti
                                     Pseudochthonius
                                             Pseudochthonius arubensis
                                             Pseudochthonius beieri
                                             Pseudochthonius billae
                                             Pseudochthonius biseriatus
                                             Pseudochthonius brasiliensis
                                             Pseudochthonius clarus
                                             Pseudochthonius congicus
                                             Pseudochthonius doctus
                                             Pseudochthonius falcatus
                                             Pseudochthonius galapagensis
                                             Pseudochthonius gracilimanus
                                             Pseudochthonius heterodentatus
                                             Pseudochthonius homodentatus
                                             Pseudochthonius insularis
                                             Pseudochthonius leleupi
                                             Pseudochthonius moralesi
                                             Pseudochthonius mundanus
                                             Pseudochthonius naranjitensis
                                             Pseudochthonius orthodactylus
                                             Pseudochthonius perreti
                                             Pseudochthonius pulchellus
                                             Pseudochthonius ricardoi
                                             Pseudochthonius simoni
                                             Pseudochthonius strinatii
                                             Pseudochthonius thibaudi
                                             Pseudochthonius troglobius
                                             Pseudochthonius tuxeni
                                             Pseudochthonius yucatanus
                                     Sathrochthoniella
                                             Sathrochthoniella zealandica
                                     Sathrochthonius
                                             Sathrochthonius crassidens
                                             Sathrochthonius insulanus
                                             Sathrochthonius kaltenbachi
                                             Sathrochthonius maoricus
                                             Sathrochthonius pefauri
                                             Sathrochthonius tuena
                                             Sathrochthonius venezuelanus
                                             Sathrochthonius webbi
                                     Spelyngochthonius
                                             Spelyngochthonius beieri
                                             Spelyngochthonius grafittii
                                             Spelyngochthonius heurtaultae
                                             Spelyngochthonius provincialis
                                             Spelyngochthonius sardous
                                     Stygiochthonius
                                             Stygiochthonius barrancoi
                                     Troglochthonius
                                             Troglochthonius doratodactylus
                                             Troglochthonius mirabilis
                                     Tyrannochthoniella
                                             Tyrannochthoniella zealandica
                                               Tyrannochthoniella zealandica foveauxana
                                               Tyrannochthoniella zealandica zealandica
                                     Tyrannochthonius
                                           [Parachthonius]
                                           [Paraliochthonius (Pholeochthonius)]
                                           [Pholeochthonius]
                                             Tyrannochthonius alabamensis
                                             Tyrannochthonius aladdinensis
                                             Tyrannochthonius albidus
                                             Tyrannochthonius amazonicus
                                             Tyrannochthonius aralu
                                             Tyrannochthonius archeri
                                             Tyrannochthonius assimilis
                                             Tyrannochthonius attenuatus
                                             Tyrannochthonius avernicola
                                             Tyrannochthonius bagus
                                             Tyrannochthonius bahamensis
                                             Tyrannochthonius barri
                                             Tyrannochthonius beieri
                                             Tyrannochthonius binoculatus
                                             Tyrannochthonius bispinosus
                                             Tyrannochthonius brasiliensis
                                             Tyrannochthonius brevimanus
                                             Tyrannochthonius brooksi
                                             Tyrannochthonius butleri
                                             Tyrannochthonius caecatus
                                             Tyrannochthonius cavernicola
                                             Tyrannochthonius cavicola
                                             Tyrannochthonius centralis
                                             Tyrannochthonius chamberlini
                                             Tyrannochthonius chamorro
                                             Tyrannochthonius charon
                                             Tyrannochthonius chelatus
                                             Tyrannochthonius confusus
                                             Tyrannochthonius contractus
                                             Tyrannochthonius convivus
                                             Tyrannochthonius curazavius
                                             Tyrannochthonius densedentatus
                                             Tyrannochthonius diabolus
                                             Tyrannochthonius ecuadoricus
                                             Tyrannochthonius elegans
                                             Tyrannochthonius erebicus
                                             Tyrannochthonius felix
                                             Tyrannochthonius fiskei
                                             Tyrannochthonius floridensis
                                             Tyrannochthonius gezei
                                             Tyrannochthonius gnomus
                                             Tyrannochthonius gomyi
                                             Tyrannochthonius grimmeti
                                             Tyrannochthonius guadeloupensis
                                             Tyrannochthonius halopotamus
                                             Tyrannochthonius helenae
                                             Tyrannochthonius heterodentatus
                                             Tyrannochthonius hoffi
                                             Tyrannochthonius horridus
                                             Tyrannochthonius hypogeus
                                             Tyrannochthonius imitatus
                                             Tyrannochthonius infernalis
                                             Tyrannochthonius innominatus
                                             Tyrannochthonius insulae
                                             Tyrannochthonius intermedius
                                             Tyrannochthonius japonicus
                                               Tyrannochthonius japonicus dogoensis
                                               Tyrannochthonius japonicus japonicus
                                             Tyrannochthonius jonesi
                                             Tyrannochthonius kermadecensis
                                             Tyrannochthonius krakatau
                                             Tyrannochthonius laevis
                                             Tyrannochthonius luxtoni
                                             Tyrannochthonius mahunkai
                                             Tyrannochthonius meneghettii
                                             Tyrannochthonius meruensis
                                             Tyrannochthonius migrans
                                             Tyrannochthonius monodi
                                             Tyrannochthonius nanus
                                             Tyrannochthonius nergal
                                             Tyrannochthonius noaensis
                                             Tyrannochthonius norfolkensis
                                             Tyrannochthonius oahuanus
                                             Tyrannochthonius oligochaetus
                                             Tyrannochthonius orpheus
                                             Tyrannochthonius osiris
                                             Tyrannochthonius ovatus
                                             Tyrannochthonius pachythorax
                                             Tyrannochthonius palauanus
                                             Tyrannochthonius pallidus
                                             Tyrannochthonius parvus
                                             Tyrannochthonius pecki
                                             Tyrannochthonius perpusillus
                                             Tyrannochthonius philippinus
                                             Tyrannochthonius pholeter
                                             Tyrannochthonius pluto
                                             Tyrannochthonius procerus
                                             Tyrannochthonius psoglavi
                                             Tyrannochthonius pupukeanus
                                             Tyrannochthonius pusillimus
                                             Tyrannochthonius pusillus
                                             Tyrannochthonius queenslandicus
                                             Tyrannochthonius rahmi
                                             Tyrannochthonius rex
                                             Tyrannochthonius riberai
                                             Tyrannochthonius robustus
                                             Tyrannochthonius rotundimanus
                                             Tyrannochthonius satan
                                             Tyrannochthonius semidentatus
                                             Tyrannochthonius semihorridus
                                             Tyrannochthonius setiger
                                             Tyrannochthonius sheltae
                                             Tyrannochthonius similidentatus
                                             Tyrannochthonius simillimus
                                             Tyrannochthonius simulans
                                             Tyrannochthonius sinensis
                                             Tyrannochthonius skeletonis
                                             Tyrannochthonius sokolovi
                                             Tyrannochthonius sparsedentatus
                                             Tyrannochthonius spinatus
                                             Tyrannochthonius steevesi
                                             Tyrannochthonius stonei
                                             Tyrannochthonius strinatii
                                             Tyrannochthonius stygius
                                             Tyrannochthonius superstes
                                             Tyrannochthonius suppressalis
                                             Tyrannochthonius swiftae
                                             Tyrannochthonius tartarus
                                             Tyrannochthonius tekauriensis
                                             Tyrannochthonius tenuis
                                             Tyrannochthonius terribilis
                                               Tyrannochthonius terribilis malaccensis
                                               Tyrannochthonius terribilis terribilis
                                             Tyrannochthonius texanus
                                             Tyrannochthonius tlilapanensis
                                             Tyrannochthonius torodei
                                             Tyrannochthonius troglobius
                                             Tyrannochthonius troglodytes
                                             Tyrannochthonius troglophilus
                                             Tyrannochthonius vampirorum
                                             Tyrannochthonius volcancillo
                                             Tyrannochthonius volcanus
                                             Tyrannochthonius wittei
                                             Tyrannochthonius wlassicsi
                                             Tyrannochthonius zonatus
                                     Vulcanochthonius
                                             Vulcanochthonius aa
                                             Vulcanochthonius howarthi
                                                   [Tyrannochthonius howarthi]
                                             Vulcanochthonius pohakuloae
                              Pseudotyrannochthoniinae
                                     Afrochthonius
                                             Afrochthonius brincki
                                             Afrochthonius ceylonicus
                                             Afrochthonius godfreyi
                                             Afrochthonius inaequalis
                                             Afrochthonius natalensis
                                             Afrochthonius reductus
                                             Afrochthonius similis
                                     Allochthonius
                                       Allochthonius (Allochthonius)
                                             Allochthonius borealis
                                             Allochthonius buanensis
                                             Allochthonius coreanus
                                             Allochthonius kinkaiensis
                                             Allochthonius montanus
                                             Allochthonius opticus
                                               Allochthonius opticus opticus
                                               Allochthonius opticus troglophilus
                                             Allochthonius shintoisticus
                                             Allochthonius tamurai
                                       Allochthonius (Urochthonius)
                                             Allochthonius biocularis
                                             Allochthonius ishikawai
                                               Allochthonius ishikawai deciclavatus
                                               Allochthonius ishikawai ishikawai
                                               Allochthonius ishikawai kyushuensis
                                               Allochthonius ishikawai shiragatakiensis
                                               Allochthonius ishikawai uenoi
                                               Allochthonius ishikawai uyamadensis
                                     Centrochthonius
                                             Centrochthonius kozlovi
                                             Centrochthonius schnitnikovi
                                             Centrochthonius sichuanensis
                                             Centrochthonius ussuriensis
                                     Pseudotyrannochthonius
                                             Pseudotyrannochthonius australiensis
                                             Pseudotyrannochthonius bornemisszai
                                             Pseudotyrannochthonius dentifer
                                             Pseudotyrannochthonius giganteus
                                             Pseudotyrannochthonius gigas
                                             Pseudotyrannochthonius gracilis
                                             Pseudotyrannochthonius hamiltonsmithi
                                             Pseudotyrannochthonius incognitus
                                             Pseudotyrannochthonius jonesi
                                             Pseudotyrannochthonius kobayashii
                                               Pseudotyrannochthonius kobayashii akiyoshiensis
                                               Pseudotyrannochthonius kobayashii dorogawaensis
                                               Pseudotyrannochthonius kobayashii kobayashii
                                             Pseudotyrannochthonius kubotai
                                             Pseudotyrannochthonius octospinosus
                                             Pseudotyrannochthonius queenslandicus
                                             Pseudotyrannochthonius rossi
                                             Pseudotyrannochthonius silvestrii
                                             Pseudotyrannochthonius solitarius
                                             Pseudotyrannochthonius tasmanicus
                                             Pseudotyrannochthonius typhlus
                                             Pseudotyrannochthonius undecimclavatus
                                               Pseudotyrannochthonius undecimclavatus kishidai
                                               Pseudotyrannochthonius undecimclavatus undecimclavatus
                                             Pseudotyrannochthonius utahensis
                                     Selachochthonius
                                           [Chthoniella]
                                           [Kafirchthonius]
                                             Selachochthonius cavernicola
                                             Selachochthonius heterodentatus
                                             Selachochthonius serratidentatus

Problematic Rooting (13589)
                                             Kewochthonius amplus
                                             Kewochthonius spingolus

Problematic Rooting (13594)
                                             Heterochthonius byzovae
                                             Heterochthonius caucasicus
                                             Heterochthonius gibbus
                                             Heterochthonius monocerus