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  Animalia
    Bilateria
     Deuterostomia
      Chordata
        Vertebrata
         Gnathostomata
          Osteichthyes
            Actinopterygii
              Neopterygii
                Teleostei
                  Acanthopterygii
                    Gasterosteiformes
                           [Pegasiformes]
                           [Syngnathiformes]
                      Gasterosteoidei
                             [Thoracostei]
                            Gasterosteidae
                                     Apeltes
                                             Apeltes quadracus
                                     Culaea
                                             Culaea inconstans
                                     Gasterostea
                                     Gasterosteus
                                           [Leiurus]
                                             Gasterosteus aculeatus
                                                   [Gasterosteus aculeatus aculeatus]
                                                   [Gasterosteus aculeatus microcephalus]
                                                   [Gasterosteus aculeatus santaannae]
                                                   [Gasterosteus aculeatus santaeannae]
                                                   [Gasterosteus aculeatus williamsoni]
                                                   [Gasterosteus argyropomus]
                                                   [Gasterosteus biaculeatus]
                                                   [Gasterosteus bispinosus]
                                                   [Gasterosteus brachycentrus]
                                                   [Gasterosteus cataphractus]
                                                   [Gasterosteus cuvieri]
                                                   [Gasterosteus gymnurus]
                                                   [Gasterosteus islandicus]
                                                   [Gasterosteus leiurus]
                                                   [Gasterosteus niger]
                                                   [Gasterosteus noveboracensis]
                                                   [Gasterosteus obolarius]
                                                   [Gasterosteus ponticus]
                                                   [Gasterosteus semiarmatus]
                                                   [Gasterosteus semiloricatus]
                                                   [Gasterosteus spinulosus]
                                                   [Gasterosteus teraculeatus]
                                                   [Gasterosteus tetracanthus]
                                                   [Gasterosteus trachurus]
                                                   [Leiurus aculeatus]
                                             Gasterosteus wheatlandi
                                     Pungitius
                                             Pungitius bussei
                                                   [Gasterosteus bussei]
                                             Pungitius hellenicus
                                             Pungitius platygaster
                                             Pungitius pungitius
                                                   [Gasterostea pungitius]
                                                   [Gasterosteus occidentalis]
                                                   [Gasterosteus pungitius]
                                                   [Pungitius pungitius occidentalis]
                                                   [Pungitius pungitius pungitius]
                                                   [Pygosteus occidentalis]
                                                   [Pygosteus pungitius]
                                             Pungitius sinensis
                                                   [Pungitius pungitius sinensis]
                                             Pungitius tymensis
                                     Pygosteus
                                     Spinachia
                                             Spinachia spinachia