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  Animalia
      Arthropoda
            Insecta
                    Coleoptera
                      Polyphaga
                        Scarabeiformia
                          Scarabaeoidea
                            Geotrupidae
                                     Australobolbus
                                             Australobolbus latipennis
                              Bolboceratinae
                                Athyreini
                                     Neoathyreus
                                             Neoathyreus fissicornis
                                             Neoathyreus mixtus
                                Bolboceratini
                                     Bolbelasmus
                                           [Kolbeus]
                                             Bolbelasmus arcuatus
                                             Bolbelasmus bajaensis
                                             Bolbelasmus hornii
                                             Bolbelasmus minor
                                             Bolbelasmus variabilis
                                     Bolbocerastes
                                             Bolbocerastes imperialis
                                               Bolbocerastes imperialis imperialis
                                               Bolbocerastes imperialis kansanus
                                             Bolbocerastes peninsularis
                                             Bolbocerastes regalis
                                             Bolbocerastes serratus
                                     Bolbocerosoma
                                       Bolbocerosoma (Bolbocerosoma)
                                             Bolbocerosoma biplagiatum
                                             Bolbocerosoma bruneri
                                             Bolbocerosoma cartwrighti
                                             Bolbocerosoma confusum
                                             Bolbocerosoma elongatum
                                             Bolbocerosoma farctum
                                             Bolbocerosoma hamatum
                                             Bolbocerosoma lepidissimum
                                             Bolbocerosoma mexicanum
                                             Bolbocerosoma pusillum
                                               Bolbocerosoma pusillum pusillum
                                               Bolbocerosoma pusillum townesi
                                             Bolbocerosoma quadricornum
                                             Bolbocerosoma ritcheri
                                             Bolbocerosoma tumefactum
                                     Bolborhombus
                                             Bolborhombus angulus
                                             Bolborhombus magnus
                                             Bolborhombus nitidus
                                             Bolborhombus parvulus
                                             Bolborhombus sallaei
                                                   [Bradycinetus carinatus]
                                     Bradycinetulus
                                           [Amechanus]
                                           [Bradycinetus]
                                             Bradycinetulus ferrugineus
                                             Bradycinetulus fossatus
                                             Bradycinetulus rex
                                     Eucanthus
                                             Eucanthus alutaceus
                                             Eucanthus greeni
                                             Eucanthus impressus
                                             Eucanthus lazarus
                                             Eucanthus mexicanus
                                             Eucanthus subtropicus
                                     Odonteus
                                             Odonteus alabamensis
                                             Odonteus cornigerus
                                             Odonteus darlingtoni
                                             Odonteus falli
                                             Odonteus filicornis
                                             Odonteus floridensis
                                             Odonteus liebecki
                                             Odonteus obesus
                                             Odonteus simi
                                             Odonteus thoracicornis
                              Geotrupinae
                                     [Lethrinae]
                                     Ceratophyus
                                             Ceratophyus gopherinus
                                     Ceratotrupes
                                             Ceratotrupes bolivari
                                     Frickius
                                     Geotrupes
                                           [Melanotrupes]
                                       Geotrupes (Anoplotrupes)
                                             Geotrupes balyi
                                             Geotrupes hornii
                                       Geotrupes (Cnemotrupes)
                                             [Onychotrupes]
                                             Geotrupes blackburnii
                                               Geotrupes blackburnii blackburnii
                                               Geotrupes blackburnii excrementi
                                             Geotrupes semiopacus
                                             Geotrupes splendidulus
                                               Geotrupes splendidulus miarophagus
                                               Geotrupes splendidulus splendidulus
                                             Geotrupes ulkei
                                       Geotrupes (Geohowdenius)
                                             Geotrupes egeriei
                                             Geotrupes opacus
                                       Geotrupes (Geotrupes)
                                             Geotrupes stercorarius
                                       Geotrupes (Megatrupes)
                                             Geotrupes cavicollis
                                             Geotrupes chevrolati
                                             Geotrupes conicollis
                                             Geotrupes gilnickii
                                             Geotrupes haldemani
                                             Geotrupes insulana
                                             Geotrupes jekelii
                                             Geotrupes lecontei
                                             Geotrupes melsheimeri
                                             Geotrupes mixtus
                                             Geotrupes ovalipennis
                                             Geotrupes similis
                                             Geotrupes starki
                                     Lethrus
                                     Mycotrupes
                                             Mycotrupes cartwrighti
                                             Mycotrupes gaigei
                                             Mycotrupes lethroides
                                             Mycotrupes pedester
                                             Mycotrupes retusus
                                                   [Geotrupes aeneus]
                                     Peltotrupes
                                             Peltotrupes profundus
                                                   [Geotrupes chalybaeus]
                                                   [Peltotrupes dubius]
                                             Peltotrupes youngi
                                     Taurocerastes
                                     Gilletinus
                                             Gilletinus variegatus