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  Animalia
    Bilateria
     Deuterostomia
      Chordata
        Vertebrata
         Gnathostomata
          Osteichthyes
            Actinopterygii
              Neopterygii
                Teleostei
                  Ostariophysi
                    Characiformes
                           [Characoidei]
                            Lebiasinidae
                              Lebiasininae
                                     Derhamia
                                             Derhamia hoffmannorum
                                     Lebiasina
                                             Lebiasina bimaculata
                                             Lebiasina chucuriensis
                                             Lebiasina floridablancaensis
                                             Lebiasina intermedia
                                             Lebiasina multimaculata
                                             Lebiasina narinensis
                                             Lebiasina provenzanoi
                                             Lebiasina uruyensis
                                             Lebiasina yuruaniensis
                                     Piabucina
                                             Piabucina astrigata
                                                   [Lebiasina astrigata]
                                             Piabucina aureoguttata
                                             Piabucina boruca
                                             Piabucina elongata
                                             Piabucina erythrinoides
                                             Piabucina festae
                                             Piabucina panamensis
                                             Piabucina pleurotaenia
                                             Piabucina unitaeniata
                              Pyrrhulininae
                                Nannostomini
                                     Nannostomus
                                           [Nannobrycon]
                                           [Poecilobrycon]
                                             Nannostomus anduzei
                                             Nannostomus beckfordi
                                                   [Nannostomus beckfordi anomalus]
                                                   [Nannostomus beckfordi aripirangensis]
                                                   [Nannostomus beckfordi beckfordi]
                                             Nannostomus bifasciatus
                                             Nannostomus britskii
                                             Nannostomus digrammus
                                             Nannostomus eques
                                                   [Nannobrycon eques]
                                                   [Poecilobrycon auratus]
                                             Nannostomus espei
                                                   [Poecilobrycon espei]
                                             Nannostomus harrisoni
                                             Nannostomus limatus
                                             Nannostomus marginatus
                                             Nannostomus marilynae
                                             Nannostomus minimus
                                             Nannostomus mortenthaleri
                                             Nannostomus nitidus
                                             Nannostomus trifasciatus
                                                   [Poecilobrycon trifasciatus]
                                             Nannostomus unifasciatus
                                                   [Nannobrycon unifasciatus]
                                                   [Nannostomus ocellatus]
                                                   [Poecilobrycon ocellatus]
                                                   [Poecilobrycon unifasciatus]
                                Pyrrhulinini
                                     Copeina
                                             Copeina guttata
                                                   [Copeina argyrops]
                                                   [Pyrrhulina guttata]
                                             Copeina osgoodi
                                     Copella
                                             Copella arnoldi
                                                   [Copeina arnoldi]
                                             Copella carsevennensis
                                             Copella compta
                                             Copella eigenmanni
                                             Copella metae
                                                   [Copeina metae]
                                             Copella nattereri
                                                   [Copeina callolepis]
                                                   [Pyrrhulina nattereri]
                                             Copella nigrofasciata
                                                   [Pyrrhulina nigrofasciata]
                                             Copella vilmae
                                     Pyrrhulina
                                             Pyrrhulina australe
                                             Pyrrhulina beni
                                             Pyrrhulina brevis
                                             Pyrrhulina eleanorae
                                             Pyrrhulina elongata
                                             Pyrrhulina filamentosa
                                             Pyrrhulina laeta
                                             Pyrrhulina lugubris
                                             Pyrrhulina macrolepis
                                             Pyrrhulina maxima
                                             Pyrrhulina melanostoma
                                             Pyrrhulina obermulleri
                                             Pyrrhulina rachoviana
                                             Pyrrhulina semifasciata
                                             Pyrrhulina spilota
                                             Pyrrhulina stoli
                                             Pyrrhulina vittata
                                             Pyrrhulina zigzag