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                            Sulidae
                                     Papasula
                                             Papasula abbotti
                                                   [Sula abbotti]
                                     Sula
                                             Sula dactylatra
                                               Sula dactylatra bedouti
                                               Sula dactylatra californica
                                               Sula dactylatra dactylatra
                                               Sula dactylatra fullagari
                                               Sula dactylatra melanops
                                               Sula dactylatra personata
                                             Sula granti
                                             Sula leucogaster
                                               Sula leucogaster brewsteri
                                               Sula leucogaster etesiaca
                                               Sula leucogaster leucogaster
                                               Sula leucogaster plotus
                                             Sula nebouxii
                                               Sula nebouxii excisa
                                               Sula nebouxii nebouxii
                                             Sula sula
                                               Sula sula rubripes
                                               Sula sula sula
                                               Sula sula websteri
                                             Sula variegata